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Vocabulary

BODC parameter semantic model biological entity development stage terms

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ID Preferred Label Definition Date
S1115 N1-6 DEPRECATED This code is equivalent to S1130 and has been deprecated. 2017-07-30
S1116 adult An animal that has reached sexual maturity 2010-02-09
S1168 aulophora larvae Organisms from the planktonic larval stage of the burrowing polychaete from the genus Lanice. 2017-06-17
S1166 auricularia The first stage of larval development of holothurians (sea cucumbers). 2017-01-24
S1117 auxospores An enlarged cell in diatoms which restores cell size following a series of divisions. 2006-03-29
S1164 blastozooids Individual organisms produced as the result of asexual reproduction such as budding 2011-08-01
S1118 calyptopis 2006-02-17
S1178 cercaria A free-swimming larval stage of trematodes (also known as parasitic flatworms or flukes) during which the parasitic fluke typically passes from an intermediate host (often a snail) to another intermediate host or to the final vertebrate host. Cercaria can develop from the sporocyst stage or from the redial stage of the trematode depending on species. 2019-08-23
S1175 cercaria purpurea The larval stage of the trematode Parorchis acanthus (urn:lsid:marinespecies.org:taxname:158687), a parasitic castrator of the common periwinkle Littorina littorea (urn:lsid:marinespecies.org:taxname:140262). Trematoda is a clade within the phylum Platyhelminthes. It includes two groups of parasitic flatworms, known as flukes. They are internal parasites of molluscs and vertebrates. 2019-08-23
S1182 chalimus The life stages of parasitic copepod species marked by their secure attachment to a host by a frontal filament. The chalimus stages follow the free-swimming copepodid stage and precede the pre-adult stages (sometimes referred to as 'sub-adult') where the organism is capable of detaching from its filament to move across the surface of its host. 2021-03-23
S115 copepodites All sub-stages of the second development stage of copepoda crustaceans following the nauplii stage (N1 to N6) and before reaching the adult stage (sometimes also labelled C6 or CVI). 2017-07-30
S111 copepodites C1 The first sub-stage of the copepodite development stage of the crustacean copepoda sub-class. 2017-07-30
S112 copepodites C1-C2 Grouping of the first 2 sub-stages of the copepodite development stage of the crustacean copepoda sub-class. 2017-07-30
S113 copepodites C1-C3 Grouping of the first 3 sub-stages of the copepodite development stage of the crustacean copepoda sub-class. 2017-07-30
S114 copepodites C1-C4 Grouping of the first, second, third and fourth sub-stages of the copepodite development stage of the crustacean copepoda sub-class. 2017-07-30
S117 copepodites C2 The second sub-stage of the copepodite development stage of the crustacean copepoda sub-class. 2017-07-30
S118 copepodites C3 The third sub-stage of the copepodite development stage of the crustacean copepoda sub-class. 2017-07-30
S119 copepodites C3-C4 Grouping of the third and fourth sub-stages of the copepodite development stage of the crustacean copepoda sub-class. 2017-07-30
S1110 copepodites C4 The fourth sub-stage of the copepodite development stage of the crustacean copepoda sub-class. 2017-07-30
S1111 copepodites C4-C5 Grouping of the fourth and fifth sub-stages of the copepodite development stage of the crustacean copepoda sub-class. 2017-07-30
S1163 copepodites C4-C5 plus adults Grouping of the last 2 sub-stages of the copepodite development stage of the crustacean copepoda sub-class plus the adult stage. 2017-07-30
S1112 copepodites C5 The fifth and last sub-stage of the copepodite development stage of the crustacean copepoda sub-class. 2017-07-30
S1114 copepodites C5 plus adults Grouping of the fifth and last sub-stages of the copepodite development stage of the crustacean copepoda sub-class plus the adult form (sometimes also labelled C6 or CVI). 2017-07-30
S116 copepodites plus adults All sub-stages of the second (copepodite) development stage of copepoda crustaceans combined with the adult stage (sometimes also labelled C6 or CVI). 2017-07-30
S1119 cyphonautes 2006-02-17
S1120 cypris The final larval stage of crustacean cirripedes or barnacles (infraclass Cirripedia) following the naupliar phase and prior to the sessile adult phase. 2017-01-24
S1121 cysts 2006-02-17
S1184 early stage eggs An embryonic development stage from extrusion of the egg to the formation of an advanced blastodisc (also known as the germinal disc or blastoderm), encompassing the zygote, morula and blastula stages of egg development. 2021-04-28
S1122 eggs Spherical or ellipsoidal bodies produced by the female of certain animals (e.g. fishes, birds, reptiles). 2017-06-19
S1179 ephyra A free-swimming precursor stage of the adult medusa stage, ephyrae (or ephyrulae) are small medusae ("medusoid heads") that separate from the polyp by budding in a process known as strobilation before metamorphosing into scyphomedusae, the life stage of the adult medusa typically identified as jellyfish. Strobils can sometimes become scyphomedusae without passing through the ephyra stage. 2019-08-29
S1181 epitoke A life stage of aquatic polychaete worms whose life cycle include a process known as "epitoky". Epitoky is the metamorphosis of a sexually immature worm (the atoke) into a sexually mature form (the epitoke) adapted for a brief free-swimming near-surface phase. 2019-11-25
S1195 eudoxid Free-living sexual unit of a majority of calycophoran siphonophores. 2021-08-24
S1189 flexion larvae A fish life stage between the start of notochord flexion and until complete notochord flexion. This is concurrent with development of the caudal-fin rays and supporting skeletal elements. 2021-04-28
S1123 furcilia Fourth free-swimming planktonic larval stage of euphausiaceans with stalked eyes and the general body form of the adult. 2010-06-21
S1156 gamete A reproductive cell (male or female) that has only half the usual number of chromosomes 2008-10-16
S1124 heterococcolithophorid 2006-02-17
S1125 holococcolithophorid 2006-02-17
S1126 hydromedusae 2006-02-17
S1171 immature This term may have subtle variations in its definition depending on the type of organism it relates to. In a general sense, it applies to organisms that are at the last stage of their juvenile development prior to a fully mature adult stage, ie when an organism looks like an adult but its reproductive organs have not yet reached maturity. 2018-05-03
S1152 indeterminate Development stage could not be reliably determined 2006-02-17
S1127 juvenile An animal that has not yet reached sexual maturity 2010-02-09
S1145 juvenile+adult Juveniles and adults have either been grouped together or cannot be reliably distinguished. 2006-02-17
S1147 juvenile+furcilia Juveniles and furcilia have either been grouped together or cannot be reliably distinguished. 2006-02-17
S1150 juvenile+sub-adult Juveniles and sub-adults have either been grouped together or cannot be reliably distinguished. 2006-02-17
S1128 larvae 2006-02-17
S1186 late stage eggs An embryonic development stage from the end of gastrulation to hatching. 2021-04-28
S1129 medusae 2006-02-17
S1167 megalopae Organisms at the final larval development stage of decapod crustaceans (e.g. crabs) following the zoea and preceding the adult stage. 2017-06-17
S1176 metacercaria A resting late larval stage of the digenetic trematodes (digeneans) with a three-host life cycle. It is a tailless stage which develops in the intermediate host either encysted in a cyst or encapsulated in a layer of host tissue. This is usually the form which is infective to the definitive host when it ingests the infected intermediate host. 2019-08-23
S1169 metanauplii Organisms from an early larval stage of some crustaceans such as krill. 2017-06-21
S1185 middle stage eggs An embryonic development stage from the beginning to the end of gastrulation. 2021-04-28
S1161 mysis Free-swimming planktonic larval forms of decapod crustaceans (especially shrimps) with three recognised development sub-stages. First sub-stage produced from protozoeae by moulting. 2010-06-21
S1130 nauplii All sub-stages of the first development stage of crustaceans after hatching. 2017-07-30
S1140 nauplii N1 The first sub-stage of the first development stage (nauplius) of crustaceans. 2017-07-30
S1154 nauplii N1-N3 Grouping of the first 3 sub-stages of the nauplius larval development stage of crustaceans. 2017-07-30
S1141 nauplii N2 The second sub-stage of the nauplius larval development stage of crustaceans. 2017-07-30
S1142 nauplii N3 The third sub-stage of the nauplius larval development stage of crustaceans. 2017-07-30
S1143 nauplii N4 The fourth sub-stage of the nauplius larval development stage of crustaceans. 2017-07-30
S1155 nauplii N4-N6 Grouping of the last 3 sub-stages of the nauplius larval development stage of crustaceans. 2017-07-30
S1144 nauplii N5 The fifth sub-stage of the nauplius larval development stage of crustaceans. 2017-07-30
S1146 nauplii N6 The sixth and last sub-stage of the nauplius larval development stage of crustaceans. 2017-07-30
S1197 nectochaeta A developmental stage found in Polychaeta species, it is a planktonic stage bearing functioning parapodia serving locomotion [Plate and Huseman 1994](https://hmr.biomedcentral.com/track/pdf/10.1007/BF02366201.pdf). 2021-08-24
S1172 newborn A newly born vertebrate (fishes, amphibians, reptiles, birds, mammals). The term 'hatchling' is used for newborn emerged from an egg. 2018-12-11
S1131 not specified No information on development stage has been provided. 2006-03-29
S1196 nurse A developmental stage in the life cycle of doliolids (Order Doliolida). The nurse stage is the terminal life stage for doliolids, often characterized by a conspicuous tail (cadophore) sprouting small trophozooids (i.e. 'nutrition zooids') also called 'gastrozooids' (i.e., feeding buds) [[Paffenhofer and Koster, 2011](https://doi.org/10.1093/plankt/fbr001); [Greer et al 2020](https://doi.org/10.1002/lno.11450)]. 2021-08-24
S1165 oozoids Individual organisms representing the asexual life form of Thaliassae (e.g. salps) and some Cnidaria; they are issued from eggs produced by sexual reproduction of blastozooids and bud asexually to produce many new sexually reproducing blastozoids. 2017-01-05
S1194 paralarva Planktonic stage of young cephalopods between hatching and sub-adult. 2021-08-24
S1132 planula The free-swimming, flattened, ciliated, bilaterally symmetrical larva of a hydrozoan cnidarian 2006-03-29
S1162 pluteus larvae Free-swimming, bilaterally symmetrical, ciliated developmental stage of some echinoderms, such as sea urchins and brittlestars 2010-12-04
S1153 post-auxospore Cell in diatoms which has restored its size by a series of divisions or is in the process of dividing following the enlarged auxospore stage. 2006-03-29
S1159 post-weaned pup A juvenile pinniped that is less than one year old and is nutritionally independent and no longer ingesting milk produced by an adult female.. 2010-03-05
S1190 postflexion larvae A fish life stage between the completion of notochord flexion and until the start of metamorphosis. Diagnostic characteristics include formation of the caudal fin and attainment of meristic complements such as fin rays. 2021-04-28
S1133 postlarvae 2006-02-17
S1151 postlarvae+juvenile Postlarvae and juveniles have either been grouped together or cannot be reliably distinguished. 2006-02-17
S1149 postlarvae+sub-adult Postlarvae and sub-adults have either been grouped together or cannot be reliably distinguished. 2006-02-17
S1191 praniza The juvenile form of a family of marine isopod crustaceans known as Gnathiidae. The adults are associated with sponges and occur in a wide range of depths. The praniza 'larva' is a temporary parasite of marine fishes. 2021-08-23
S1183 pre-adult and adult motile stages The pre-adult and adult life stages of parasitic copepod species, commonly referred to as 'sea lice', that are marked by the ability of parasites to move freely across the surface of their host or swim freely in the water column. Sea lice motile stages are preceded by the chalimus stages in which the organism is attached to its host by a frontal filament and exclude the free-swimming naupliar and copepodid stages. 2021-03-23
S1158 pre-weaned pup A juvenile pinniped that is less than one year old and is ingesting milk produced by an adult female. 2010-03-05
S1188 preflexion larvae A fish life stage between complete yolk-sac absorption and until the start of notochord flexion. 2021-04-28
S1160 protozoeae Free-swimming planktonic larval forms of decapod crustaceans with three recognised development sub-stages (Dall et al., 1990). 2010-06-21
S1157 pup A juvenile pinniped that is less than one year old 2010-02-09
S1180 pupa An organism at the pupal stage. A life stage of some insects undergoing a complete metamorphosis between immature (larva) and mature (adult or imago) stages. 2019-11-15
S1134 resting spores 2006-02-17
S1174 scyphistoma The fixed segmented polyp-like stage in the life cycle of a jellyfish, which reproduces asexually by budding (strobilation). 2019-08-23
S1173 smolt A young salmon or sea-going trout that is at the stage of development when it assumes a silvery color and is physiologically ready to migrate to the sea. 2019-07-16
S1170 spat Shellfish larvae of the Bivalvia class (e.g. oysters, mussels, scallops) that have settled on a substrate following the end of their planktonic life stage (plural form may be spat or spats). 2018-03-26
S1135 spores 2006-02-17
S1177 sporocyst A term used for a development stage in the lifecycle of Apicomplexa parasites and trematode flatworm parasites (flukes). In the trematodes lifecycle it is an elongated sac-like structure that is formed in the initial stage of infection of an intermediate host (typically a snail). It develops from a miracidium and evolve into a redia. 2019-08-23
S120 sub-adult 2006-02-17
S1148 sub-adult+adult Sub-adults and adults have either been grouped together or cannot be reliably distinguished. 2006-02-17
S1192 trochophore Free-swimming planktonic marine larva occurring in the first developmental stage of Molluscs and certain marine annelids. The trochophore is the developmental stage prior to veliger. 2021-08-24
S1136 vegetative cell 2006-02-17
S1193 veliger Planktonic larva typical of many marine and freshwater snails and bivalves. The veliger develops from the trochophore larva. 2021-08-24
S1187 yolk-sac larvae A fish life stage between hatching and until complete absorption of yolk-sac. 2021-04-28
S1137 zoeae Free-swimming planktonic larval form of decapod crustaceans (especially crabs) with a relatively large cephalothorax, conspicuous eyes, and fringed antennae and mouthparts. 2010-06-21
S1138 zooids 2006-02-17
S1139 zoospores 2006-02-17